We appreciate the very helpful comments of P. Garfinkel, J. Hadam, S. Pethybridge, PHI Reviewers, and members of the 2006 course in Ecology and Epidemiology of Plant Pathogens at KSU. It’s also a pleasure to acknowledge support by the U.S. National Science Foundation under Grants DEB-0130692, DEB-0516046, EF-0525712 (as part of the joint NSF-NIH Ecology of Infectious Disease program) and DBI-0630726, by the Ecological Genomics Initiative of Kansas through NSF Grant No. EPS-0236913 with matching funds from the Kansas Technology Enterprise Corporation, by the Office of Science (Program in Ecosystem Research), U.S. Department of Energy, Grant No. DE-FG02-04ER63892, by the U.S. Agency for International Development for the Sustainable Agriculture and Natural Resources Management Collaborative Research Support Program (SANREM CRSP) under terms of Cooperative Agreement Award No. EPP-A-00-04-00013-00 to the Office of International Research and Development at Virginia Tech and for the Integrated Pest Management CRSP, by USDA grant 2002-34103-11746, and by the NSF Long Term Ecological Research Program at Konza Prairie. This is Kansas State Experiment Station Contribution No. 07-310-J.